A Revision of the collection of Noctuidae (Lepidoptera)

reported from Korea by O. Herz in 1904

Vladimir S. Kononenko

Institute of Biology and Pedology, Far Eastern Branch of the Russian Academy of Sciences, RF-690022 Vladivostok, Russia

Abstract The collection of Noctuidae reported by Herz (1904) from Korean the peninsula is revised and its corrected list is given. The identity of some species is confirmed or corrected with comments related to recent Korean faunistic literatures. The types of the noctuids described by Herz (1905) are revised and the lectotypes for seven taxa described are designated. New synonymies for two taxa (Coryta grisea Leech, 1900 = Polydesma striata Herz, 1904, syn. n., Apopestes indica Moore = Apopestes spectrum var. coreana Herz, 1904, syn. n.) are proposed. The male genitalia of types and authentical specimens for the taxa described by Herz are illustrated.

Key words Lepidoptera, Noctuidae, revision of collection, nomenclature, Korean peninsula

Introduction

A well-known Russian lepidopterist, the conservator of Russian Imperial Zoological Museum, Otto Fedrovich Herz (1852-1905) was one of the first researcher of the Lepidoptera fauna in the Korean peninsula. He collected Lepidoptera in Korea by himself during the summer season 1884 at a small villiage Pung-Tung near Kimwah (38.5﹉N, 128﹉E), Kangwon province. Herz also examined some materials collected by M. Jankowsky in 1894 at a valley of Tumankang river in the northern part of Korea. As a results of his study for the Korean Lepidoptera, Herz published ※The list of Noctuidae and Geometridae from Korea§ (Herz, 1904). It was a complete and accurate check list at that time, with numerous additions to previous publications (Leech, 1889, 1900) and with descriptions of nine new species, a variety, and a new genus of the Noctuidae.

Herz﹊s materials was a part of collection of Great Prinz Nikolai Mikhailovich Romanov - an uncle of the last Russian Imperior Nikolai Romanov, famoust lepidoptera amateur, sponsor of many entomological expeditions and projects, a publisher of famous Russian lepidopterological issue ※Memoirs sur Lepidopterous§. Collection of N. M. Romanov was donated by him to Zoolological Museum of Imperior Academy of Sciences in St.- Petersburg in 1900. At present these materials including Herz collection from Korea are preserved in the Zoological Institute of Russian Academy of Sciences. Like other members of Imperior﹊s family N. M. Romanov was shuted after revolution by bolsheviks in 1917 in Petrograd.

Since Herz﹊s publication, during the subsequent 1890 years, the nomenclature and taxonomic status of many taxa for the Korean noctuids have been changed considerably. The present paper is based on the examination of Herz's materials and type specimens described by him from Korea, and dividing in three parts:

1) A revised list of Herz﹊s material. In this part, the names of taxa are cited as same order and nomenclature as used by Herz in the left column. In the right column, the names of each taxon are arranged by the correct identification under the current status and nomenclature where they have been changed. The following abbreviations are used in the parenthesis: misid. - misidentification; missp. - misspelling; mispl. - misplacing, for the species treated in another genus (or under another generic name) in current sense; syn. - synonymic name at present, hom. - homonym; emend. - unnecessary emendation. Some taxa required more complete explanation are marked by indexes.

2) Comments. Comments to Herz﹊s list contains explanations for some critical species corresponding to names used by him. In most cases, comments are given in relation to using Herz﹊s data in Korean literatures on Noctuidae.

3) A revised catalogue of the types described by Herz. The taxa described by Herz`s are indicated in the left column of Herz﹊s list with asterisk (*). In the catalogue, the names of species-group taxa are arranged in the alphabetical order with the following standard data: name of taxon, originally accepted taxonomic category (in round bracket): (sp) - species, (var) - variety; original combination with author and year of reference to the original description; type locality (TL); type material (TM). The data of labels are quoted exactly, separated by a stroke (/). The synonyms and current status for each taxa are given under the paragraph ※ID§ (identity) with references to corresponding literary sources.

The specimens in Herz﹊ collection have small, printed standard labels with black frame: ※Korea§, on upper side ※Herz§ or ※Korea Herz 1884§ or ※Korea§ and ※Jankowsky 1894§. In addition all specimens have the labels in Russian: ※洵眇看看. 派盆洎. 炫洩洵眇看癸竿洫洩祉癸相看眇皇洩祇癸.§ which means ※Collection of Great Prince Nicolai Mikhailovich [Romanov]※. The specimens from type series are supplied by blue or pinky label with ※Origin§. Some type specimens illustrated in photo plates in Herz﹊s publication have the pins with upper part cut. Under description of a new of taxa Herz gave the label ※Origin§ to all specimens in type series, but did not designate as ※Type§ for a single specimen . Therefore the lectotypes are selected for seven taxa. In three cases the taxa were described, based on a single specimen, which is the holotype according to International Code of Zoological Nomenclature (ICNZ). For illustrations of genitalia, both types and authentical specimens identified by comparing with types were used.

A revised list of Herz﹊s material

Original names by Herz (1904) Corrections (Current name)

Diphtera niveola Motsh. (mispl.) - Belciades niveola (Motsch., 1866)

Diphtera virens Butl. (misid., mispl.) - Belciana staudingeri (Leech, 1900)

Diphtera alpium Osb. var. murrhina Graes. - Moma kolthoffi (Bryk, 1948)

(misid., mispl.)

Aplectoides caliginea Butl. (mispl.) - Anacronicta caliginea (Butl., 1881)

Acronicta major Brem. - Acronicta major Brem., 1861

Acronicta strigosa F. - Acronicta strigosa (Den. & Schiff., 1775)

Acronicta albonigra Herz *(mispl.) - Hampsonidia albonigra (Herz, 1904)

Acronicta jankowskii Obth. (mispl.) - Hampsonidia jankowskii (Obth., 1880)

Acronicta tridens D.& S. var. increta Butl. (syn.) - Acronicta intermedia (Warr., 1909 )

Acronicta cuspis Hb. var. leucocuspis Butl. - Acronicta leucocuspis (Butl., 1878)

Acronicta raphaelis Obth. (emend.) - Acronicta raphael (Obth., 1884)

Acronicta digna Butl. - Acronicta digna (Butl., 1881)

Acronicta rumicis L. - Acronicta rumicis (L., 1758)

Acronicta carbonaria Graes. - Acronicta carbonaria (Graes., [1898] 1888

Acronicta lutea Brem. - Acronicta lutea (Brem. & Grey, 1852)

Agrotis obscura Brahm. (syn., mispl.) - Spaelotis ravida (Den. & Schiff., 1775)

Agrotis fuscostigma Brem. (mispl.) - Xestia fuscostigma (Brem., 1861)

Agrotis exusta Butl. (mispl.) - Sineugraphe exusta (Butl., 1878)

Agrotis exusta, ab. nigromaculata Graes. - Sineugraphe bipartita (Graes., [1889] 1888)

(syn., mispl.)

Agrotis baja F. (misid., mispl.) - Xestia tabida (Butl., 1878)

Agrotis kollari Led. (mispl.) - Xestia kollari (Led., 1853)

Agrotis cissigma Men. (mispl.) - Lycophotia cissigma (Men., 1859)

Agrotis stupenda Butl. (mispl.) - Xestia stupenda (Butl., 1878)

Agrotis triangularis Moore (mispl.) - Albocosta triangularis (Moore, 1867)

Agrotis islandica Stgr var. rossica Stgr. (mispl.) - Euxoa ochrogaster rossica (Stgr., 1881)

Agrotis nigricostata Stgr. (syn., mispl.) - Agrotis tokionis Butl., 1881

Agrotis ypsilon Rott. (emend., miss. auth.) - Agrotis ipsilon (Hfn., 1766)

Agrotis segetum D.& S. - Agrotis segetum (Den. & Schiff, 1775.

Agrotis sibirica Bsdv. - Euxoa sibirica (Bsdv., 1832)

Mamestra adjuncta Stgr. (syn., mispl.) - Polia bombycina grisea (Butl., 1878)

Mamestra brassicae L. - Mamestra brassicae (L., 1758)

Mamestra splendes Hb. (mispl.) - Lacanobia splendes (Hb., [1808])

Dianthoecia cucubali Fluess. (misid., mispl.) - Sideridis honei (Yoshimoto, 1989)

Miana askoldis Obth. (mispl.) - Leucapamea askoldis (Obth., 1880)

Miana segregata Butl. (mispl.) - Niphonyx segregata (Butl., 1878 )

Miana variegata Obth.(syn., mispl.) - Pyrrhidivalva sordida (Butl., 1881)

Miana strigilis Cl. var. an ab. (mispl.) - ※Oligia§ leuconephra (Hmps., 1808 )

Luperina depravata Butl. (mispl.) - Spodoptera depravata (Butl., 1879)

Hadena subpulchra Alph. (mispl.) - Euromoia subpulchra (Alph., 1897)

Hadena confecta Walk. (misid., misspl.) - Olivinebula oberthueri (Stgr., 1892)

Hadena funerea Hein. (syn., mispl.) - Apamea aquila oriens (Warr., 1911)

Hadena muscosa Butl. (syn.) - Atrachea nitens (Butl., 1878)

Hadena jankowskii Obth. (mispl.) - Atrachea jankowskii (Obth., 1879)

Hadena arcta Led. (part., mispl.) - Eucarta arcta (Led., 1853)

Hadena arcta Led. (part., misid., mispl.) - Eucarta fasciata (Butl., 1878)

Hadena arctides Stgr. (mispl.) - Eucarta arctides (Stgr., 1888)

Dipterygia scabriuscula L. (misid.) - Dipterygia caliginosa (Walk., 1858)

Rhisogramma saga Butl. (mispl.) - Auchmis saga (Butl., 1878)

Antha grata Butl. - Antha grata (Butl., 1881)

Chloantha intermedia Brem. (emend., mispl.) - Actinotia intermediata (Brem., 1861)

Callopistria purpureofasciata Piller (syn.) - Callopistria juventina (Stoll., 1782)

Callopistria miracula Herz* (mispl.) - Callopistria miracula Herz, 1904 (mispl.)

Callopistria argyrosticta Butl. - Callopistria argyrosticta (Butl., 1881)

Trachea atriplicis L. - Trachea atriplicis (L., 1758)

Trachea melanospila Koll. var. lucia Butl. (hom.) - Trachea punkikonis Mats. , 1928 (lucia Butl.,

nec Feld. & Rghr., 1874)

Trachea kosakka Obth. (syn.) - Trachea melanospila Koll., [1844]

Euplexia lucipara L. - Euplexia lucipara (L., 1758)

Mania muscivirens Butl. - Mormo muscivirens Butl., 1878

Orthogonia sera Feld. - Orthogonia sera Feld. & Feld., 1862

Naenia contaminata Walk. - Naenia contaminata (Walk., 1865)

Helotropha leucostigma Hb. var. laevis Butl. (mispl.) - Celaena leucostigma (Hb., [1808])

Virgo amoena Stgr. (syn.) - Virgo datanidia (Butl., 1885)

Nonagria graminea Graes. (mispl.) - Xylomoia graminea (Graes., [1889] 1888)

Sesamia inferens Walk. - Sesamia inferens (Walk., 1856)

Leucania impuncta Stgr. (hom.mispl.) - Mythimna simplex (Leech, 1900), repl. name.

Leucania pallens var. melania Stgr. (misident, - Mythimna pallens (L., 1758)

mispl.)

Leucania velutina Ev. (mispl.) - Mythimna velutina (Ev., 1846)

Leucania radiata Brem. (mispl.) - Mythimna radiata (Brem., 1861)

Leucania albicosta Moore (hom. misid., mispl.) - Mythimna separata (Walk., 1865)

Leucania unipuncta Haw. (misid. mispl.) - Mythimna separata (Walk., 1865)

Leucania turca L. (mispl.) - Mythimna turca (L., 1758)

Leucania divergens Butl. (mispl.) - Mythimna divergens Butl., 1887

Leucania biundulata Motsch. (syn. uncert.) - Mythimna grandis Butl., 1878

Leucania rufipennis Butl. (mispl.) - Mythimna rufipennis Butl., 1878

Caradrina albosignata Obth. et ab. caeca Obth. - Athetis albisignatha (Obth., 1879)

(mispl.)

Caradrina lenta Tr. var. lentina Stgr. (syn.) - Athetis furvula (Hb., [1808])

Hydrilla gluteosa Tr. (mispl.) - Athetis gluteosa (Tr., 1835)

Hydrilla maurella Stgr. (mispl.) - Stygiodrina maurella (Stgr., 1888)

Hydrilla illustrata Stgr. (mispl.) - Condica illustrata (Stgr., 1888)

Hydrilla tristis Brem. ab. lugens Stgr. (syn.) - Athetis lepigone (Mshl., 1907)

Amphipyra livida F. var. corvina Motsch. - Amphipyra livida corvina Motsch. , 1866

Amphipyra jankowskii Obth. - Amphipyra jankowskii Obth., 1884

Amphipyra erebina Butl. - Amphipyra erebina Butl., 1878

Amphipyra pyramidea L. var. monolitha Gn. - ?Amphipyra monolitha Gn., 1852 (id. uncert)

Eutelia geyeri F. & R. - Eutelia geyeri Feld. & Rog., 1774

Anuga constricta Gn. (misid.) - Anuga multiplicans Walk., 1858

Erastria argentula Hb. var. amurula Stgr. - Deltote bankiana (F., 1775)

(syn., mispl.)

Erastria obliterata Rmbr. (mispl.) - Phyllophila obliterata (Rmbr., 1833)

Erastria pusilla View. (syn.) - Pseudeustrotia candidula (D. & S.)

Erastria nemorum Obth. (mispl.) - Deltote nemorum (Obth., 1880)

Erastria fasciana L. var. guenei Fall. (syn.) - Protodeltote pydarga (Hfn., 1766)

Erastria fasciana L. var. stygia Butl. (mispl.) - ?Sugia stygia (Butl., 1878)

Erastria fasciana L. var. distinguenda Stgr.(mispl.) - Protodeltote distinguenda (Stgr., 1888)

Erastria fentoni Butl. var. versicolor Obth. (mispl.) - Erastroides fentoni (Obth., 1881)

Rivula sericealis Scop. - Rivula sericealis (Scop., 1763)

Milichia gemella Leech (mispl.) - Dysmilichia gemella (Leech, 1889)

Ilattia stellata Butl. (mispl.) - Amyna stellata (Butl., 1878)

Amyna octo Gn. - Amyna octo Gn., 1852

Emmelia trabealis Scop. - Emmelia trabealis (Scop.,1763)

Tarache bicoloria Leech (mispl.) - Acontia bicolora (Leech, 1889)

Scoliopteryx libatrix L. - Scoliopteryx libatrix (L., 1758)

Calpe capucina Esp. (syn.) - Calyptra thalictri (Bkh., 1790)

Calpe aureola Graes. (syn.) - Calyptra lata (Butl., 1881)

Plusiodonta casta Butl. - Plusiodonta casta Butl., 1878

Telesilla amethystina Hb. (mispl.) - Eucarta amethystina (Hb., [1803])

Telesilla virgo Tr. (mispl.) - Eucarta virgo (Tr., 1835)

Plusia agnata Stgr. (mispl.) - Acanthoplusia agnata (Stgr., 1892)

Plusia ornatissima Walk. (misid., mispl.) - Antoculeora locuples (Obth., 1880)

Thermesia ussuriensis Brem. (mispl.) - Blasticorhinus ussuriensis (Brem., 1861)

Remigia annetta Butl. (mispl.) - Mocis annetta (Butl., 1878)

Remigia archesia Cram. (syn.) - Mocis undata (F., 1775)

Pelamia electaria Brem. (mispl.) - Melapia electaria (Brem., 1864)

Polydesma striata Herz* (syn. n.) - Cortyta grisea Leech, 1900

Zethes musculus Men. (mispl.) - Arytrura musculus (Men., 1852)

Zethes subfalcata Men. (mispl.) - Arytrura subfalcata (Men., 1852)

Ophiusa obscura Brem. (mispl.) - Dysgonia obscura (Brem. & Grey, 1853)

Ophiusa coreana Leech (mispl.) - Dysgonia coreana (Leech, 1889)

Ophiusa arcuata Moore var. mandshuriana Stgr. - Dysgonia mandshuriana (Stgr., 1892)

(mispl.)

Ophiusa annalis Gn. (misid., mispl.) - Dysgonia arctotaenia Gn., 1852

Ophiusa dulcis Butl. (mispl.) - Dysgonia dulcis (Butl., 1878)

Ophiusa dotata F. (mispl.) - Artena dotata (F., 1794)

Spirama rectifasciata Men. (mispl.) - Metopta rectifasciata (Men., 1863)

Sypna picta Butl. (mispl.) - Sypnoides picta (Butl., 1877)

Sypna rectifasciata Graes. (syn., mispl.) - Sypnoides hercules (Butl., 1881)

Edesena hamada Feld. - Edesena hamada (Feld. & Rog., 1874)

Pseudophila amata Brem & Grey (mispl.) - Chrysorithrum amatum (Brem. & Grey, 1853)

Pseudophila flavomaculata Brem. (mispl.) - Chrysorithrum flavomaculatum (Brem., 1861)

Arcte coerulea Gn. (missp.) - Arcte coerula (Gn., 1852)

Lagoptera juno Dalm. (mispl.) - Thyas juno (Dalm. , 1823)

Ophideres tyrannus Gn. (mispl.) - Adris tyrannus (Gn., 1852)

Catocala electa Bkh. - Catocala electa Viev., 1790

Catocala nupta L. var. obscurata Obth. - Catocala nupta (L., 1761)

Catocala fulminea Scop. - Catocala fulminea (Scop., 1763)

Catocala koreana Stgr. - Catocala koreana Stgr., 1892

Catocala abamita Brem. & Grey - Catocala abamita Brem. & Grey, 1853

Catocala agitatrix Graes. - Catocala agitatrix Graes., [1889] 1888

Catocala obscena Alph. - Catocala obscena Alph., 1879

Catocala helena Ev. - Catocala helena Ev., 1856

Catocala doerriesi Stgr. - Catocala doerriesi Stgr., 1888

Catocala obliterata Men. (syn.) - Catocala praegnax Walk., [1858] 1859

Catocala puella Leech. - Catocala puella Leech., 1889

Koraia pirata Herz* - Catocala pirata (Herz, 1904)

Apopestes spectrum L. var. koreana Herz* (syn.) - Apopestes indica (Moore, 1882)

Apopestes cataphanes Hb. - Autophila inconspicua Butl., 1881

var. inconspiqua Butl. (mispl.)

Eccrita maxima Brem. (mispl.) - Lygephila maxima (Brem., 1861)

Toxocampa recta Brem. (mispl.) - Lygephila recta (Brem., 1864)

Toxocampa moellendorffi Herz* (mispl.) - Lygephila moellendorffi (Herz, 1904)

Aventiola pusilla Butl. - Aventiola pusilla (Butl., 1879)

Egnasia pulcherima Butl. (mispl.) - Lophoruza pulcherrima (Butl., 1879)

Epizeuxis lunulata Herz * (syn.) - Idia curvipalpis (Butl., 1879)

Capnodes lignea Butl. (mispl.) - Anatatha lignea (Butl., 1879)

Capnodes cremata Butl. (mispl.) - Diomea cremata (Butl., 1878)

Capnodes cinerea Butl. (mispl.) - Hypostrotia cinerea (Butl., 1878)

Bleptina spacoalis Walk. (mispl.) - Bertula spacoalis (Walk., 1859)

Simplicia rectalis Ev. - Simplicia rectalis (Ev., 1842)

Gisira signata Butl. (mispl.) - Cidariplura signata (Butl., 1879)

Zanclognatha tarsiplumalis Hb. - Zanclognatha lunalis (Scop., 1763)

Zanclognatha tarsipennalis Tr. - Zanclognatha tarsipennalis (Tr., 1835)

Zanclognatha tarsicrinalis Knoch. - Herminia tarsicrinalis (Knoch., 1782)

Zanclognatha fumosa Butl. - Zanclognatha fumosa Butl., 1879

Zanclognatha stramentacealis Brem. (mispl.) - Herminia stramentacealis Brem., 1864

Zanclognatha umbrosalis Stgr. - Zanclognatha umbrosalis Stgr., 1892

Zanclognatha trilinealis Brem. - Paracolax trilinealis (Brem., 1864)

Zanclognatha bistrigata Stgr. (mispl.) - Bertula bistrigata (Stgr., 1888)

Zanclognatha punctalis Herz*(mispl.) - Sinarella punctalis (Herz, 1904)

Pangrapta vasava Butl. - Pangrapta vasava Butl., 1881

Pangrapta flavomacula Stgr. - Pangrapta flavomacula Stgr., 1888

Pangrapta suaveola Stgr. - Pangrapta suaveola Stgr., 1888

Pangrapta indentalis Leech - Pangrapta indentalis Leech, 1888

Egnasia costaemacula Stgr. (mispl.) - Pangrapta costaemacula Stgr., 1888

Marmorinia obscurata Butl. (mispl.) - Pangrapta obscurata (Butl., 1888)

Hydrillodes morosa Butl. - Hydrillodes morosa Butl., 1879

Dierna timandra Alph. (mispl.) - Naganoella timandra (Alph., 1897)

Gynaephila maculifera Stgr. - Gynaephila maculifera Stgr., 1892

Madopa salicalis D. & S. (mispl.) - Colobochyla salicalis (Den. & Schiff., 1775)

Madopa flavomacula Obth. (mispl.) - Paragabara flavomacula (Obth., 1888)

Herminia gryphalis H.-S. (mispl.) - Polypogon gryphalis (H.-S., 1851)

Herminia derivalis Hb. (syn., mispl.) - Paracolax tristalis (F., 1794)

Herminia tentacularia Hb. (mispl.) - Polypogon tentacularia (Hb., 1758)

Bomolocha nigrobasalis Herz* (mispl.) - Hypena nigrobasalis (Herz, 1904))

Bomolocha stygiana Butl. (mispl.) - Hypena stygiana (Butl., 1878)

Bomolocha bicoloralis Graes. (mispl.) - Hypena bicoloralis (Graes. , [1889] 1888)

Dichromia amica Butl. (mispl.) - Hypena amica (Butl., 1878)

Hypena proboscidalis L. var. deleta Stgr. - Hypena proboscidalis (L., 1758)

Hypena tristalis Led. - Hypena tristalis Led., 1853

Hypenodes separatalis Herz*(mispl.) - Schrankia separatalis (Herz, 1904)

Comments

Diphtera alpium var. murrhina.- The species was misidentified by Herz as Diphtera alpium var. murrhina (Graes.), but it was described later as Moma kolthoffi (Bryk, 1948) (= fulvicollis de Lattin, 1948). For the cited synonymy, see Kononenko, 1996. The name Moma orion var. murrhina Graeser, [1889] 1888 is a junior synonym of Moma alpium (Osbeck, 1778).

Agrotis baja - At present, Xestia baja ([Denis & Schifferm ller], 1775) and Xestia tabida are treated as two distinct species, forming a sister group since Boursin (1963). The former is distributed in the Western Palaearctic region, and the Far Eastern population belongs to the latter. However this specific pair needs a revision for clarifying their status as two distinct species or a subspecies. Author identified Korean population as X. tabida.

Agrotis islandica var. rossica - In the current treatment, Euxoa islandica (Staudinger, 1857) is considered as a synonym of the Holarctic species Euxoa ochrogaster (Guen e, 1953) (Lafontaine, 1987). In the Western Palaearctic region, the latter represented as a subspecies Euxoa ochrogaster ssp. islandica (Staudinger, 1857) and in the Eastern Palaearctic region as Euxoa ochrogaster ssp. rossica (Staudinger, 1881).

Dianthoecia cucubali - The species reported as Dianthoecia cucubali (a junior subjective synonym of Sideridis (Aneda) rivularis (Fabricius, 1775)) by Herz was reidentified as Sideridis (Aneda) honeyi Yoshimoto, 1989. Since Herz, this species had been reported in Korean publications as Hadena rivularis, H. rivularis mandarina (Leech, 1900), H. cucubali ([Denis & Schifferm뚴ler], 1775) or H. cucubali mandarina. It was listes to the Check List of Insects from Korea (1994) as Hadena rivularis. Yoshimoto (1989) separated Hadena mandarina (Lech, 1900) from rivularis as a distinct species and described subspecies H. rivularis honeyi. The taxonomic status of the last was rised by Hacker (1996) to a good species. S. rivularis, S. honeyi and S. mandarina represent as a close sister species-group, which are hardly identifiable by the external appearance. Only one species, S. honeyi has been found in Korea, although all three species are distributed in the south of Russian Far East and N China (Hacker, 1996); the occurrence of other species in Korea is not impossible, but need a further investigation.

Miana strigilis var. an ab. - The taxon reported as Miana strigilis Cl. var. an ab.[Oligia strigilis (Linnaeus,1758)] by Herz was re-identified as ※Oligia§ leuconephra. Oligia strigilis is a Western Palaearctic species and not distribute in the Far East. Since Herz, it was not reported for Korea or neibouring countries. The generic position of ※Oligia§ leuconephra is still unclear.

Hadena subpulchra - The species was described from Korea by Alpheraky (1897) based on specimen collected by Jankowski from N Korea and reported for Korea by Leech (1900). Herz (1904) reported it as Hadena subpulchra. After these records it was referred under the genus Hadena in Korean literatures. Kononenko (1976) transferred Hadena subpulchra to the genus Euromoia (Subfam. Acronictinae) and reidentified Euromoia subpulchra as E. mixta Staudinger, 1892 by Kozhanchikov (1950). The species was also incorrectly listed in the Check List of Insects from Korea (1994) under combinations Hadena subpulchra and also Euromoia subpulchra.

Hadena confecta - The species, reported by Herz [ZI, St. Petersburg] as Hadena confecta, was reidentified as Olivinebula oberthueri (Staudinger, 1892). This species was also misidentified by Leech (1900) and reported for Korea as Triphaenopsis pulcherrima Moore. Both species, Olivenebula pulcherrima (Moore, 1867) and Olivenebula confecta (Walker,1858) are distinct from O. oberthueri and are not represented in the Far East. Both names, pulcherrima and confecta were incorrectly used in Korean literatures. The name Hadena confecta was incorrectly spelled by Pak (1959) as conferta. This name as well as pulcherrima was listed in the Check List of Insects from Korea (1994) as ※Hadena conferta [sic, misspelling] Walker§ and Triphaenopsis pulcherima [sic] Moore. All three names (pulcherrima, confecta and conferta) related to Olivinebula oberthueri and should be excluded from the faunal list of the Korean Noctuidae.

Hadena jankowskii - The species was reported as Hadena jankowskii by Herz (1904). Since Herz, it has been incorrectly referred in Korean literatures as Hadena jankowskii or as Trachea jankowskii Obth. Under the last combination as well as Atrachea jankowskii Oberth뚼 it was listed in the Check List of Insects from Korea (1994). The species was also incorrectly illustrated in Nam et al. (1982: Pl. 42, fig. 295a) as Atrachea nitens (Butler, 1878).

Hadena arcta - The material reported as Hadena arcta by Herz was reidentified as Eucarta arcta (Lederer, 1853) and partially as Eucarta fasciata (Butler, 1878).

Dipterygia scabriuscula - Herz misidentified Korean specimen of Dipterygia caliginosa as the Western Palaearctic species, Dipterygia scabriuscula. Since Herz, this name has been cited in some Korean publications. The former species is not distributed in the Far East.

Trachea melanospila - The species reported as Trachea melanospila var. lucia by Herz is currently considered as Trachea punkikonis Matsumura, 1927, which is a replaced name for the secondary homonym, Hadena lucia Butler, 1878. Since Herz, it has been incorrectly referred in the Korean literature as Trachea auriplena (Walker, 1857) or as T. lucilla [sic] Butler [T. lucia Butler, 1878]. It was incorrectly listed in the Check List of Insects from Korea (1994) as Trachea auriplena, althouth this Indian-Himalayan species is not distributed in the Far East.

Leucania albicosta and Leucania unipuncta - The species repoted as Leucania unipuncta and Leucania albicosta by Herz are the misidentification of Mythimna (Pseudaletia) separata Walker, 1865. The former species do not distributed in the Far East. Mythimna (Pseudaletia) pallidicosta (Hampson, 1894) is a replacement name for Aletia albicosta Moore, 1882 (homonym) applied to a distinct species. However, under the examination of Herzs material from Korea [ZIN, Spb.], this species was not found. Some specimens of M. separata (reported by Herz as Leucania unipuncta) have some reddish tint in the coloration of forewings, which make them somewhat similar to M. pallidicosta, but it lies in the limits of specific variability of M. separata. The name albicosta probably after Herz (1904) was repeated in Korean literature (Pak, 1959; Nam et al., 1982) and listed in the Check List of Insects from Korea (1994) under a homonym name of Pseudaletia albicosta. The species M. pallidicosta (albicosta, hom.) is distributed in South East Asia, and migrating moths reaching to South of Japan. It was not found in Korea. Herzs record is probably due to the misidentification.

Amphipyra pyramidea var. monolitha - The identity of Korean and continental Far Eastern population of Amphipyra pyramidea/ monolitha with N Indian species Amphipyra monolitha Guenee, 1852 is still unclear. A further examination for this taxon is needed.

Anuga constricta. - The species reported for Korea as Anuga constricta Guenee, 1852 by Herz is a misidentification of Anuga multiplicans (Walker). Herz incorrectly placed the name multiplicans in the synonymy of constricta. However these two taxa currently are treated as distinct species (Holloway, 1985; Poole, 1989). A. constricta is distributed in SE Asia, north to Vietnam and Thailand, and A. multiplicans has more wide distribution and reaches to China, Japan and Korea. Under this name the species was reported for Korea by Pak (1964: 54) and also by Park and Lee (1977: 5). Both names, Anuga multiplicans and Anuga constricta were listed in the Check List of Insects from Korea (1994). However, the name Anuga constricta should be excluded from Korean faunal list.

Erastria fasciana var. stygia - The material reported by Herz was examined and reidentified as P. pygarga. The name stygia is currently applied to a distinct species Sugia stygia (Butler, 1878). It hardly distinguishable from very polymorphic P. pygarga by the external appearance,. The species Sugia stygia was reported in Korean literatures (Maruta 1929a; 1929b), probably on the base of earlier reports by Leech (1889; 1900) and Herz (1904). The occurrence of Sugia stygia is possible in Korea, but needs a further investigation.

Plusia ornatissima - The records of Antoculeora ornatissima (Walker, 1858) by Herz and other authors from Korea and the Far East are due to misidentification of Antoculeora locuples (Oberthr, 1881). These taxa currently are considered as two distinct species (Kitching, 1987, Kononenko, 1990; Ronkay, 1996). A. ornatissima is a Indian-Himalayan species, and not distributed in the Far East. All records of A. ornatissima from the Far East may belong to A. locuples.

Ophiusa annalis - The species reported for Korea (Herz, 1904: 43) as Ophiusa analis Gn. [Dysgonia analis (Guenee, 1852)] by Herz is a misidentification of Dysgonia stuposa (Fabricius, 1794). Material reported by Herz was examined and reidentified as D. stuposa. The name D. analis is incorrectly listed in the Check List of Insects from Korea (1994).

Egnasia costaemacula - The identity of this species from Herzs collection is uncertain. The species Pangrapta costaemacula (Staudinger, 1888) forms a sister species-group with its allies recently described as P. duplex (Sugi & Kononenko, 1996). The latter is the most common species of Pangrapta in Korea, Japan and China. During the examination of Herz`s material, the specimen identified as costaemacula was not dissected, however, the occurrence of costaemacula in Korea, is quite possible in its northern part at least.

Sypna rectifasciata - Material reported as Sypna rectifasciata by Herz was reidentified as Sypnoides hercules (Butler, 1881). The name rectifasciata is a junior synonym of hercules. Both name is listed in the Check List of Insects from Korea (1994) as Sypna rectifasciata and Sypnoides hercules. The former should be excluded from the faunal list of Korean Noctuidae as a synonym.

Catocala puella Leech - Author could not find specimens in the collection of ZI, St. Petersburg, which was collected from Korea and identified by Herz. The species Catocala puella Leech, 1889, described from Korea, is known only by the type, preserved in BMNH, London, and also by two documentated records in N and S Korea.

A revised catalogue of the types

albonigra (sp) - Acronicta albonigra Herz, 1904: 269, pl.1, fig. 3. TL: Korea, Pun-tung villiage [38.5* N, 128* E]. TM: Lectotype (hereby designated): male with labels: ※Korea/Herz/Orig./洵眇看看. 派盆- 洎. 炫洩洵眇看癸竿洫洩祉癸相看眇皇洩祇癸§. Genit. prep. N 17749. Paralectotypes: 6 males with the same labels, one male with label ※Prep. 3487§.

ID: Hampsonia albonigra (Herz, 1904), a good species (Subf. Acronictinae).

Remarks. A distinct species, distributed in Korea, Japan, Russian Far East. In current treatment it is placed to the genus Craniophora (Sugi, 1892). However, author treats Hampsonia (type species Apatele jankowskii Oberth뚼, 1880) and Craniophora (type species Noctua ligustri [Denis & Schifferm뚴ler], 1775) as a distinct genera. Male genitalia for the authentical specimen of Hampsonia albonigra (Herz, 1904) is illustrated in the fig. 1.

koreana (var) - Apopestes spectrum L. var. koreana Herz, 1904: 315, pl.1, fig. 11. TL: North Korea [?Tumangang valley]. TM: Lectotype (hereby designated): female with label: ※Jankowsky, 1894 /Korea/ Orig./洵眇看看. 派盆洎. 炫洩洵眇看癸竿洫洩祉癸相看眇皇洩祇癸§. The pin of the lectotype is cut at upper part. The specimen was illustrated by Herz. Paralectotypes: 5﹡, 5＋, with same labels; 1＋: ※Korea / Herz / Orig./洵眇看看. 派盆洎. 炫洩洵眇看癸竿§ and red label ※Sp. Spectrum L. var phantasma Ev.§

ID: Apopestes indica (Moore, 1883) (=Apopestes spectrum L. var. koreana Herz, syn. n.) (Subf. Ophiderinae).

Remarks. The species was described by Herz from North Korea as Apopestes spectrum var. koreana Herz. Its status was revised by Boursin (1955) to a good species. Later Boursin put the name koreana to a synonymy of Apopestes indica Moore, 1883 in his unpublished Check List of palaearctic noctuidae, but it was not officially synonymized. The types of Apopestes indica are conspecific with koreana (L. Ronkay, pers. comm.). Poole (1989) incorrectly referred Apopestes indica as a synonym of Apopestes spectrum (Esper, 1787) and koreana as a good species. Both names Apopestes koreana and Apopestes spectrum were incorrectly included in the Check List of Insects from Korea (1994). The species A. spectrun does not distributed in the Far East, the name koreana is a synonym of Apopestes indica. The genitalia of the authentical specimen, compared with types koreana, is illustrated in the fig. 2.

miracula (sp) - Callopistria miracula Herz, 1904: 284, pl.1, fig. 10. TL: Korea, Pun-tung villiage [38.5* N, 128* E]. TM: Holotype: a male with labels: ※Korea / Herz /Orig./洵眇看看. 派盆洎. 炫洩洵眇看癸竿洫洩祉-癸相看眇皇洩祇癸.§

ID: ※Callopistria§ miracula Herz, 1904, a good species (Subf. Amphipyrinae).

Remarks. A distinct specimen was found in the recent Korean collection. Its generic position is uncertain. Herz placed it to Callopistria, however, its external morphology and male genitalia do not corrrespond to Callopistria juventina (Stoll, 1782), the type species of the genus. Apparently this species belongs to lower Amphipyrinae. Until now the species is known only from Korea. Male genitalia of the holotype of Callopistria miracula Herz, 1904, is illustrated in the fig. 3.

moellendorffi (sp) - Toxocampa moellendorffi Herz, 1904: 318, pl.1, fig. 1. TL: Korea, Pun-tung williage [38.5* N, 128* E]. TM: Lectotype (hereby designated): male, ※Korea / Herz / Orig./洵眇看看. 派盆洎. 炫洩洵眇看癸竿洫洩祉癸相看眇皇洩祇癸§. The specimen was illustrated by Herz, the upper part of pin is cut. Paralectotype: 1﹡ with same labels.

ID: Lygephila moellendorffi (Herz, 1904), a good species (Subf. Ophiderinae).

Remarks. A good species, however, no additional record after Herz's description is appeared. Male genitalia of the paralectotype is illustrated in the fig. 4.

lunulata (sp) - Epizeuxis (Helia) lunulata Herz, 1904: 320, pl.1, fig. 12. TL: Korea, Pun-tung villiage [38.5* N, 128* E]. TM: Lectotype (hereby designated): a male, ※Korea / Herz / Orig. [red label]/洵眇看看. 派盆洎. 炫洩洵眇看癸竿洫洩祉癸相看眇皇洩祇癸§. Paralectotypes: 1﹡ (without abdomen) with same label.

ID: Idia curvipalpis (Butler 1879) (Subf. Herminiinae).

Remarks. The name is currently considered to be a junior synonym of Idia curvipalpis (Butler), synonymized by Remm & Martin (1979). The species was illustrated by Owada (1982). Male genitalia of the authentical specimen is illustrated in the fig. 5.

nigrobasalis (sp) - Bomolocha nigrobasalis Herz, 1904: 329, pl.1, fig. 13. TL: Korea, Pun-tung villiage [38.5* N, 128* E]. TM: Lectotype (hereby designated): a male, ※Korea Herz 1884 / Orig./洵眇看看. 派盆洎. 炫洩洵眇看癸竿洫洩祉癸相看眇皇洩祇癸,§ genitalia preparate No 17490. Paralectotypes: 2﹡, with labels: ※Jankowsky, 1894 /Korea/ Orig./洵眇看看. 派盆洎. 炫洩洵眇看癸竿洫洩祉癸相看眇皇洩祇癸§.

ID: Hypena nigrobasalis (Herz, 1904), a good species (Subf. Hypeninae).

Remarks. Currently considered as a good species. The species was illustrated by Sugi (1982). Male genitalia of the lectotype is illustrated in the fig. 6.

pirata (sp) - Koraia pirata Herz, 1904: 314, pl.1, fig. 18. TL: Korea, Pun-tung villiage [38.5﹉N, 128﹉E]. TM: Lectotype (hereby designated): 1＋ with lables: ※Korea / Herz / Orig./洵眇看看. 派盆洎. 炫洩洵眇-看癸竿洫洩祉癸相看眇皇洩祇癸§. The pin of the lectotype is cut at upper part. The specimen was illustrated by Herz. Paralectotype: a female with the same labels and additional label ※Micropreparate N 14652§.

ID: Catocala pirata (Herz, 1904), a good species (Subf. Catocalinae).

Remarks. The genus Koraia Herz described by Herz (1904) on the same publication was synonymised with Catocala Schrank by Kononenko (1990: 29, note 86). Male genitalia of the authentical specimen to compare with types is illustrated in the fig. 7.

punctalis (sp) - Zanclognatha punctalis Herz, 1904: 324, pl.1, fig. 9. TL: Korea, Pun-tung villiage [38.5﹉N, 128﹉E]. TM: Holotype: a female with labels: ※Korea / Herz / Orig./洵眇看看. 派盆洎. 炫洩洵眇-看癸竿洫洩祉癸相看眇皇洩祇癸§.

ID: Sinarella punctalis (Herz, 1904) (Subf. Herminiinae).

Remarks. Currently considered as a good species. It was illustrated by Owada (1982; 1987). Male genitalia was illustrated by Yamamoto & Sugi (1955) as Zanclognatha nigrobasalis Yamamoto & Sugi (junior synonym of punctalis), Remm & Martin (1987) and also by Owada. Male genitalia of the authentical specimen is illustrated in the fig. 8.

separatalis (sp) - Hypenodes separatalis Herz, 1904: 331, pl.1, fig. 7. TL: Korea, Pun-tung williage [38.5﹉N, 128﹉E]. TM: Lectotype (hereby designated): a male with labels ※Korea / Herz / Orig./洵眇看看. 派盆洎. 炫洩洵眇看癸竿洫洩祉癸相看眇皇洩祇癸.§ Paralectotypes: 2﹡, 1＋, with the same labels.

ID: Schrankia separatalis (Herz, 1904) (Subf. Hypenodinae).

Remarks. Currently considered as a good species. It was illustrated by Sugi (1982). Male genitalia (fig. 9) was illustrated by Remm & Martin (1979).

striata (sp) - Polydesma striata Herz, 1904: 303, pl.1, fig. 4. TL: North Korea [?Tumanghan valley]. TM: Holotype: male with labels: ※Jankowsky 1894/ Orig./洵眇看看. 派盆洎. 炫洩洵眇看癸竿洫洩祉癸相看眇皇洩祇-癸§.

ID: Polydesma striata Herz, 1904 is a junior synonym (syn. n.) of Cortyta grisea (Leech, 1900) (Subf. Catocalinae).

Remarks. The type specimen is identical with Cortyta grisea (Leech, 1900, described from China (Trans. ent. Soc. Lond. 1900: 552, Polydesma). The latter taxon was illustrated by Hampson (1913, text fig. 82) and Chen (1982). The new synonymy was established, based on the comparision of imagos and male genitalia of the holotype of Polydesma striata Herz with specimen from Korea which was identified as Cortyta grisea (Leech, 1900). In the original description, Herz incorrectly indicated the type as a female but the holotype of P. striata is a male. The specimen is in some worn condition, therefore wing pattern is not contrasted as fresh ones. Male genitalia of the holotype of Polygesma striata Herz is illustrated in the fig. 10.

Acknowledgments

I wish to express my cordial thanks to Dr. A. L. Lvovsky and Dr. I. L. Sukhareva, Zoological Institute, Russian Academy of Sciences, St.-Peterburg for their kind consultation and help during my study, with supplying the collection under their curation and preparation of the present article. I am deeply grateful to Dr. S. B. Ahn (Agricultural Sciences and Technology Institute, Rural Development Administration, Suwon, Korea) and Prof. K. T. Park (Center for Insect Systematics, Chuncheon, Korea) for their critical review of the manuscript.

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Herz (1904)縑曖ボフ奩紫縑憮爾堅脹嫘釭寞峞曖營薑葬

V.S. Kononenko

楝衛嬴婁ピ錳斛翕碟模儅僭饜曄翱掘模, RF-690000 綰塭蛤爾蝶驚, 楝衛嬴

1904喇 Herz縑曖ボフ奩紫縑憮爾堅脹嫘釭寞峞蹗菟擊營薑葬ビ堅, 熱薑フピ貲曖蹗跡煙擊爾堅フ棻. 傳フ斜縑曖ボ閬婞攄戲煎嫦ォ腎歷湍蹗菟縑渠ボ撲貲婁ヘ眷營翕薑唸婁蒂薑葬ビ艘戲貊, 橾睡蹗曖賅衝ォ獄薑葬諦ヘ眷蹗縑渠フ Lectotype擊撲薑ビ艘棻. 舒謙, striata Herz諦 coreana Herz擊陝陝 Coryta grisea Leech諦 Apopestes indica Moore曖 synonym戲煎薑葬ビ堅, 賃賃輿蹂蹗縑渠ボ撩醱曖餌霞婁熱飄儅衝晦蒂紫ボビ艘棻.

匐儀橫 : 釭綠跡, 嫘釭寞婁, 營薑葬, 貲貲敘擒, フ奩紫

(Received: November 5, 1997)

(Accepted: September 10, 1998)